Professor
Ph.D. Evolutionary Zoology, 1967, University of Texas
M.A. Evolutionary Zoology, 1964, University of Texas
B.A. Biology (Honors), 1962, University of Louisville
Teaching and research in evolutionary and behavioral ecology; resource control and reproductive success in vertebrates, including humans; integration of evolutionary theory and resource management; resources and reproductive variance; reproductive and resource tradeoffs for modern women.

